What does :- mean in Emoticons?

This page is about the meanings of the acronym/abbreviation/shorthand :- in the Internet field in general and in the Emoticons terminology in particular.

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What does :- mean?

an animal that produces gametes (spermatozoa) that can fertilize female gametes (ova)

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":-." Abbreviations.com. STANDS4 LLC, 2015. Web. 27 Feb. 2015. <http://www.abbreviations.com/term/14386>.

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